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It is possible to play an Atari 2600 using a Sega Genesis controller. The reason why the Sega Genesis controllers work on the Atari 2600 is because both the Genesis and the 2600 use DE9 ports. These ports were a popular type of port, and were used for the Sega Megadrive, the Atari 7200, Commodore 64, MSX, and the 3DO.

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